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Within the open ocean—particularly within the deep-water column, in deep sediments, and in deep crustal environments—carbon and power sources are extraordinarily scarce (oligotrophic), however active microorganisms are present (though their progress charges could also be extremely low). 58:755-805; Rothschild, L.J., and Mancinelli, R.L., 2001, Life in extreme environments, Nature 409:1092-1101; Silver, S., and Phung, L.T., 1996, Bacterial heavy metal resistance: New surprises, Annu. In a single, the consequences of extremes in pressure and temperature lengthen into the cytoplasm, and intracellular biosynthesis, metabolism, and macromolecular buildings are tailored to operate underneath such conditions.

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